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JASON PIERS WILTON HALL
Department of Entomology
National Museum of Natural History
Smithsonian Institution
Washington, DC, USA
Tel: (202) 633 1047
Email: halljrios@yahoo.com

Education:
1994-1999

Ph.D., Department of Entomology and Nematology,
University of Florida, Gainesville, USA

1990-1993 B.A., M.A. Honours, Department of Zoology,
St. Johns College, Oxford University, Oxford, UK

Grants and Awards:
2007-2011 National Science Foundation Grant
(Principal Investigator on Biodiversity Surveys & Inventories Grant DEB# 0639977 - "Collaborative Research: The Butterflies of Ecuador (Lepidoptera: Papilionoidea) - A Comprehensive Survey of a Megadiverse Fauna")
2003

Royal Entomological Society Award
(for best paper in the journal Systematic Entomology during 2001-2: Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for speciation mechanisms in the Andes)

2001-2006 National Science Foundation Grant
(Principal Investigator on Biodiversity Surveys & Inventories Grant # 0103746 - "Survey of the Andean butterfly fauna of Ecuador")
2001-2002Smithsonian Postdoctoral Fellowship
(National Museum of Natural History, Washington, DC)
1999-2000Smithsonian Postdoctoral Fellowship
(National Museum of Natural History, Washington, DC)
1999J. Turner Predoctoral Fellowship
(University of Florida, Gainesville, FL)
1998Collections Studies Grant
(American Museum of Natural History, New York, NY)
1997-2000National Geographic Society Grant
(Joint Principal Investigator on Research and Exploration Grant # 96-5751)
1996Sigma-Xi, Grant-in-aid of Research
1995Sigma-Xi, Grant-in-aid of Research
1993

Grants awarded for "Oxford and Cambridge Butterfly Expedition to Ecuador 1993"
(including Royal Entomological Society, Oxford University [Poulton Fund], and Cambridge University [Balfour Browne Fund])

Positions held:
2005President of the Entomological Society of Washington, Washington, DC
2004President-elect of the Entomological Society of Washington, Washington, DC
2002-

Research Entomologist, Department of Entomology, National Museum of Natural History, Smithsonian Institution (Trust Fund)

1999-2002

Smithsonian Postdoctoral Fellow, Department of Entomology, National Museum of Natural History, Smithsonian Institution

1999-2002

Research Associate, Department of Entomology, National Museum of Natural History, Smithsonian Institution

1999-2000

Affiliate Faculty, Department of Zoology, University of Florida

1994-1999

Research Assistant, Schaus Swallowtail Recovery Program, University of Florida

Research interests:

Broadly, my research interests cover the evolution, systematics, phylogenetics, biogeography, ecology and conservation of butterflies, particularly those from the Neotropics and those belonging to the family Riodinidae. The systematics of this almost exclusively Neotropical family of approximately 1400 species was until recently perhaps the most poorly known of all the butterfly families. As a group it also presents an endlessly fascinating array of ecological research topics, most of which are in their infancy, including species specific male perching behaviors in time and space which relate to reproductive isolation and niche partitioning, mimicry amongst each other and of almost every other butterfly family, and myrmecophily and larval-ant communication. My research on riodinid butterflies has shed light on several topics of broad evolutionary interest, including the evolution of myrmecophily and aphytophagy, providing new theories on the relationships between morphology and nutritional ecology, generating more detailed hypotheses of Amazonian area relationships and vertical parapatric montane speciation than have ever been published before using either vertebrates or invertebrates, and a revised estimate of the age of the Riodinidae based on evidence from Dominican amber fossils.

Specific short-term research goals include creating natural generic classifications for the most poorly known riodinid tribes, the Nymphidiini (the largest myrmecophilous riodinid group) and the Symmachiini, using predominantly morphological character sets but also molecular ones when the former do not provide sufficient phylogenetic information, and generating a comprehensive generic-level phylogenetic hypothesis for the Riodinidae based on molecular evidence from 2-4 genes (in collaboration with Naomi Pierce's Harvard lab). My involvement with the alpha taxonomy of butterflies, particularly riodinids, will continue as an essential component of completing two longer term research goals, a five volume comprehensive work on the Papilionoidea of Ecuador (with Keith R. Willmott), the first for any Neotropical country, and a book on the Riodinidae of the Neotropics, providing a comprehensive systematic and pictorial treatment, and a synthesis of our biological knowledge on the group.

During the course of my research, I have also visited numerous private collections around the World, and most of the major museums throughout the Americas and Europe to examine, dissect and curate Neotropical butterflies:

   Americas: Cambridge, USA - MCZ; New York, USA - AMNH; Reading, USA - RPM; Pittsburgh, USA - CMNH; Washington, USA - USNM; Gainesville, USA - FSCA; Sarasota, USA - AME; Quito, Ecuador - MECN and QCAZ; Lima, Peru - MUSM; and
   Europe: London, UK - BMNH; Oxford, UK - OXF; Paris, France - MNHN; Berlin, Germany - ZMHU; Dresden, Germany - SMTD; Frankfurt, Germany - SMF; Munich, Germany - ZSM; Stuttgart, Germany - SMS.

Field work:
I have conducted fieldwork in east Africa, throughout Asia and Australasia, and the majority of countries in Central and tropical South America as follows:

   Neotropics: Mexico (3 trips), Guatemala, Belize, Honduras, Costa Rica, Venezuela, Trinidad & Tobago, Ecuador (12 trips), Peru, Bolivia, Brazil;
   Afrotropics: Kenya;
   Oriental region: Malaysia, Taiwan, Philippines (2 trips), Papua New Guinea, Australia, Solomon Islands.

Since 1991, I have accomplished nearly a year and a half of intensive fieldwork in Ecuador, where, with a colleague (Keith R. Willmott), I have gathered voluminous ecological information and compiled one of the most extensive butterfly research collections and databases for any Neotropical country.

Scientific Publications:
Books:

1999a.

Hall, J. P. W. A Revision of the Genus Theope: its Systematics and Biology (Lepidoptera: Riodinidae: Nymphidiini). Scientific Publishers, Gainesville. vii + 127pp.

2005a

Hall, J. P. W. A Phylogenetic Revision of the Napaeina (Lepidoptera: Riodinidae: Mesosemiini). Entomological Society of Washington, Washington, DC. 235 pp.


2009a.

Hall, J. P. W. A Monograph of the Nymphidiina: Phylogeny, Taxonomy, Biology and Biogeography (Lepidoptera: Riodinidae: Nymphidiini). The Entomological Society of Washington, Washington, DC. c. 500 pp. [in prep.]

201X.

Hall, J. P. W., and K. R. Willmott. The Butterflies of Ecuador and their Natural History. Part III: Riodinidae. Oxford University Press, Oxford, UK. [in prep.]

201X.

Hall, J. P. W., K. R. Willmott, and R. C. Busby. The Butterflies of Ecuador and their Natural History. Part IV: Lycaenidae. Oxford University Press, Oxford, UK. [in prep.]

201X.

Willmott, K. R., and J. P. W. Hall. The Butterflies of Ecuador and their Natural History. Part I: Papilionidae, Pieridae and Nymphalidae (Libytheinae, Danainae, Nymphalinae, Heliconinae, Limenitidinae). Oxford University Press, Oxford, UK. [in prep.]

201X.

Willmott, K. R., and J. P. W. Hall. The Butterflies of Ecuador and their Natural History. Part II: Nymphalidae (Biblidinae, Apaturinae, Charaxinae, Satyrinae, Morphinae). Oxford University Press, Oxford, UK. [in prep.]

201X.

Willmott, K. R., and J. P. W. Hall. The Butterflies of Ecuador. A Field Guide. Oxford University Press, Oxford, UK. [in prep.]

Papers: [PDF = electronic pdf file available]
1993.

Hall, J. P. W., and K. R. Willmott. Temporal and elevational additions of distribution for Costa Rican butterflies. Tropical Lepidoptera, 4(2): 139-141. PDF

1994

Willmott, K. R., and J. P. W. Hall. Four new species of riodinids from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 5(2): 87-91. PDF

1995a.

Hall, J. P. W., and K. R. Willmott. Two new species of Mesene from western Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 110-112. PDF

1995b.

Hall, J. P. W., and K. R. Willmott. Five new species and a new genus of riodinid from the cloud forests of eastern Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 131-135. PDF

1995c.

Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, with descriptions of five new species (Lepidoptera: Riodinidae). Tropical Lepidoptera, 6(2): 136-143. PDF

1995d.

Willmott, K. R., and J. P. W. Hall. Two new species of satyrines from Ecuador (Nymphalidae: Satyrinae). Tropical Lepidoptera, 6(2): 103-105. PDF

1995e.

Willmott, K. R., and J. P. W. Hall. Two new species of Adelpha from eastern Ecuador (Lepidoptera: Nymphalidae). Tropical Lepidoptera, 6(2): 106-109. PDF

1996a.

Hall, J. P. W., and K. R. Willmott. The genus Theope: four new species and a new subspecies (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 63-67. PDF

1996b.

Hall, J. P. W., and K. R. Willmott. Notes on the genus Argyrogrammana, Part 2, with one new species. Tropical Lepidoptera, 7(1): 71-80. PDF

1996c.

Hall, J. P. W., and K. R. Willmott. Systematics of the riodinid tribe Symmachiini, with the description of a new genus and five new species from Ecuador, Venezuela and Brazil (Lepidoptera: Riodinidae). Lambillionea, 96:00:00 637-660. PDF

1996d.

DeVries, P. J., and J. P. W. Hall. Two new species of riodinids from Costa Rica (Lepidoptera: Riodinidae). Tropical Lepidoptera, 7(1): 87-90. PDF

1996e.

Torres, R., J. P. W. Hall, K. R. Willmott, and K. Johnson. A new genus of elfin butterflies from the northern high Andes (Lepidoptera: Lycaenidae). Tropical Lepidoptera, 7(1): 81-86. PDF

1997a.

Hall, J. P. W., and G. T. Austin. Riodinidae of Rondônia, Brazil: a new species of Theope (Lepidoptera: Riodinidae). Tropical Lepidoptera, 8(2): 101-102. PDF

1997b.

Hall, J. P. W., and K. R. Willmott. The Butterflies of Venezuela. Part I. Nymphalidae I (Limenitidinae, Apaturinae, Charaxinae). A Comprehensive Guide to the Identification of Adult Nymphalidae, Ppailionidae and Pieridae. by Andrew F. E. Nield. Tropical Lepidoptera, 8(1): 46 [book review] PDF

1998a.

Hall, J. P. W. A review of the genus Sarota (Lepidoptera: Riodinidae), pp. 1-21. In: J. P. W. Hall (ed.) A Contribution to Riodinid Systematics. Tropical Lepidoptera, 9(2), Supplement 1 Download reprint [PDF 1] [PDF 2]

1998b.

Hall, J. P. W. Six new species in the "foliorum group" of Theope (Lepidoptera: Riodinidae). Lambillionea, 98(4): 562-573. PDF

1998c.

Hall, J. P. W., and K. R. Willmott. Four new riodinid species from eastern Ecuador (Lepidoptera: Riodinidae). Lambillionea, 98:00:00 325-334. PDF

1998d.

Hall, J. P. W., and K. R. Willmott. Nine new species and one new subspecies of Euselasia from Ecuador (Lepidoptera: Riodinidae), pp. 27-35. In: J. P. W. Hall (ed.) A Contribution to Riodinid Systematics. Tropical Lepidoptera, 9(2), Supplement 1 PDF

1998e.

Hall, J. P. W., and K. R. Willmott. Three new species of Riodinini from the cloud forests of Ecuador (Lepidoptera: Riodinidae), pp. 22-26. In: J. P. W. Hall (ed.) A Contribution to Riodinid Systematics. Tropical Lepidoptera, 9(2), Supplement 1 PDF

1998f.

Hall, J. P. W., and D. J. Harvey. A new genus of riodinid, with a new species from Panama (Lepidoptera: Riodinidae: Nymphidiini), pp. 36-41. In: J. P. W. Hall (ed.) A Contribution to Riodinid Systematics. Tropical Lepidoptera, 9(2), Supplement 1 PDF

1999a.

Hall, J. P. W.The Genus Theope and Relatives: their Systematics and Biology (Lepidoptera: Riodinidae: Nymphidiini). Gainesville: University of Florida. Ph.D. Dissertation. viii + 381pp.

1999b.

Hall, J. P. W., and E. Furtado. Two new species and one subspecies of riodinid from southwest Brazil (Lepidoptera: Riodinidae). Tropical Lepidoptera, 10(1): 19-24. PDF

1999c.

Hall, J. P. W., and J. B. Heppner. Lemoniadini, a corrected tribal name in the Riodininae (Lepidoptera: Riodinidae). Tropical Lepidoptera, 10(1): 30 PDF

1999d.

Pyrcz, T. W., K. R. Willmott, and J. P. W. Hall. Contribution to the knowledge of Ecuadorian Pronophilini. Part III. Three new species and five new subspecies of Lymanopoda (Lepidoptera: Nymphalidae: Satyrinae). Genus, 10(3): 497-522. Download reprint [PDF 1] [PDF 2]

1999e.

Willmott, K. R., and J. P. W. Hall. Taxonomic notes on Ecuadorian Adelpha with the description of two new species and seven new subspecies (Lepidoptera: Nymphalidae: Limenitidinae). Tropical Lepidoptera, 10(1): 1-17. Download reprint [PDF 1] [PDF 2]

2000a.

Hall, J. P. W. Two new genera in the Neotropical riodinid tribe Nymphidiini (Riodinidae). Journal of the Lepidopterists' Society, 54(2): 41-46. PDF

2000b.

Hall, J. P. W., and K. R. Willmott. Patterns of feeding behaviour in adult male riodinid butterflies and their relationship to morphology and ecology. Biological Journal of the Linnean Society, 69(1): 1-23. PDF

2001a.

Hall, J. P. W. A revision of the new riodinid butterfly genus Dachetola (Lepidoptera: Riodinidae: Riodinini). Journal of the New York Entomological Society,109(2): 183-195. PDF

2001b.

Hall, J. P. W., and D. J. Harvey. A phylogenetic analysis of the Neotropical riodinid butterfly genera Juditha, Lemonias, Thisbe, and Uraneis, with a revision of Juditha (Lepidoptera: Riodinidae: Nymphidiini). Systematic Entomology, 26(4): 453-490. PDF

2001c.

Hall, J. P. W., and D. J. Harvey. A phylogenetic revision of the Charis gynaea group (Lepidoptera: Riodinidae) with comments on historical relationships among Neotropical areas of endemism. Annals of the Entomological Society of America, 94(5): 631-647. PDF

2001d.

Hall, J. P. W., and D. J. Harvey. A reassessment of Calociasma with the description of a new genus and a new species (Lepidoptera: Riodinidae: Nymphidiini). Journal of the New York Entomological Society, 109(2): 196-205. PDF

2001e.

Hall, J. P. W., and G. Lamas. Five new riodinid species from northwestern dry forest and northeastern Andean cloud forest habitats in Peru (Lepidoptera: Riodinidae). Revista Peruana Entomologia, 42:00:00 9-19. PDF

2001f.

Willmott, K. R., J. P. W. Hall, and G. Lamas. Systematics of Hypanartia (Lepidoptera: Nymphalidae: Nymphalinae), with a test for speciation mechanisms in the Andes. Systematic Entomology, 26(4): 369-399. PDF

2001g.

Willmott, K. R., L. M. Constantino, and J. P. W. Hall. A review of Colobura (Lepidoptera: Nymphalidae: Limenitidinae), with description of a new species and illustration of the immature stages. Annals of the Entomological Society of America, 94(2): 185-196. PDF

2002a.

Hall, J. P. W. A review of the new riodinid butterfly genus Panaropsis (Lepidoptera: Riodinidae: Symmachiini). Proceedings of the Entomological Society of Washington, 104(1): 63-72. PDF

2002b.

Hall, J. P. W. A review of Chalodeta with a revision of the Chalodeta chaonitis group(Lepidoptera: Riodinidae). Proceedings of the Entomological Society of Washington, 104(2): 376-389. PDF

2002c.

Hall, J. P. W. Phylogeny of the riodinid butterfly subtribe Theopeina (Lepidoptera: Riodinidae: Nymphidiini). Systematic Entomology, 27(2): 139-168. PDF

2002d.

Hall, J. P. W. A phylogenetic revision of Calydna and relatives (Lepidoptera: Riodinidae). Insect Systematics and Evolution, 33(2): 185-237.

2002e.

Hall, J. P. W., and D. J. Harvey. A survey of androconial organs in the Riodinidae (Lepidoptera). Zoological Journal of the Linnean Society, 136:00:00 171-197. PDF

2002f.

Hall, J. P. W., and D. J. Harvey. The phylogeography of Amazonia revisited: new evidence from riodinid butterflies. Evolution, 56(7): 1489-1497. PDF

2002g.

Hall, J. P. W., and D. J. Harvey. A phylogenetic review of Charis and Calephelis (Lepidoptera: Riodinidae). Annals of the Entomological Society of America, 95(4): 407-421. PDF

2002h.

Hall, J. P. W., and D. J. Harvey. A revision of the Neotropical butterfly genus Seco Hall and Harvey (Lepidoptera: Riodinidae). Proceedings of the Entomological Society of Washington, 104(4): 941-947. PDF

2002i.

Hall, J. P. W., and D. J. Harvey. Basal subtribes of the Nymphidiini (Lepidoptera: Riodinidae): phylogeny and myrmecophily. Cladistics, 18(6): 539-569.

2002j.

Harvey, D. J., and J. P. W. Hall. Phylogenetic revision of the Charis cleonus complex (Lepidoptera: Riodinidae). Systematic Entomology, 27(3): 265-300. PDF

2003a.

Hall, J. P. W. Phylogenetic reassessment of the five forewing radial-veined tribes of the Riodininae (Lepidoptera: Riodinidae). Systematic Entomology, 28(1): 23-37. PDF

2003b.

Hall, J. P. W., and C. J. Callaghan. A revision of the new riodinid butterfly genus Pseudotinea (Lepidoptera: Riodinidae). Journal of Natural History, 37(7): 821-837. PDF

2004a.

Hall, J. P. W.The Genus Adelpha: its Systematics, Biology and Biogeography (Lepidoptera: Nymphalidae: Limenitidini). by Keith R. Willmott. Proceedings of the Entomological Society of Washington, 106(3): 728-730. [book review] PDF

2004b.

Hall, J. P. W. Metalmark Butterflies (Lepidoptera: Riodinidae), pp. 1383-1386. In: J. L. Capinera (ed.) Encyclopedia of Entomology, Vol. 2. Kluwer Academic Publishers, Dordrecht, Netherlands. PDF

2004c.

Hall, J. P. W., and D. J. Harvey. Five new riodinid species from the Napo region of Ecuador and Peru (Lepidoptera: Riodinidae). Tropical Lepidoptera,12(1-2)("2001"): 35-41. PDF

2004e.

Hall, J. P. W., D. J. Harvey, and D. H. Janzen. Life history of Calydna sturnula with a review of larval and pupal balloon setae in the Riodinidae. Annals of the Entomological Society of America, 97(2): 310-321. PDF

2004f.

Hall, J. P. W., and G. Lamas. A new cloud forest species of Calydna (Lepidoptera: Riodinidae) from Peru, with a revised phylogeny of the C. hiria group. Proceedings of the Entomological Society of Washington, 106(4): 733-738. PDF

2004g.

Hall, J. P. W., R. K. Robbins, and D. J. Harvey. Extinction and biogeography in the Caribbean: new evidence from a fossil riodinid butterfly in Dominican amber. Proceedings of the Royal Society of London B, 271(1541): 797-801. PDF

2004h.

Cock, M. J. W., and J. P. W. Hall. Immature stages of Calydna venusta morio (Riodinidae)from Trinidad. Journal of the Lepidopterists' Society, 58(1): 51-53. PDF

2004i.

Willmott, K. R., and J. P. W. Hall. Taxonomic notes on the genus Zaretis, with the description of a new species (Lepidoptera: Nymphalidae: Charaxinae). Tropical Lepidoptera, 12(1-2)("2001"): 29-34. PDF

2005a.

Hall, J. P. W. A review of the Metacharis syloes group (Lepidoptera: Riodinidae), with the description of two new west Andean species. Proceedings of the Entomological Society of Washington, 107(1): 200-208. PDF

2005b.

Hall, J. P. W. Montane speciation patterns in Ithomiola butterflies (Lepidoptera: Riodinidae): are they consistently moving up in the World? Proceedings of the Royal Society of London B, 272(1580): 2457-2466. PDF

2005c.

Hall, J. P. W., and K. R. Willmott. A new species of Paiwarria (Lepidoptera: Lycaenidae: Eumaeini) from western Ecuador. Proceedings of the Entomological Society of Washington, 107(4): 960-967. PDF

2005d.

Hall, J. P. W., K. R. Willmott and R. C. Busby. Five new Penaincisalia species (Lepidoptera: Lycaenidae: Eumaeini) from the Andes of southern Ecuador and northern Peru. Zootaxa, 797: 1-20. PDF

2006a.

Hall, J. P. W. A remarkable new riodinid species, Stalachtis halloweeni (Riodinidae: Stalachtini), from Mount Ayanganna, Guyana. Journal of the Lepidopterists' Society, 60(3): 138-142.

2006b.

Pyrcz, T. W., K. R. Willmott, J. P. W. Hall and A. L. Viloria. A review of the genus Manerebia Staudinger (Lepidoptera: Nymphalidae: Satyrinae) in the northern Andes. Journal of Research on the Lepidoptera, 39: 37-79. PDF

2007a.

Hall, J. P. W. Phylogenetic revision of the new riodinid butterfly genus Minstrellus (Lepidoptera: Riodinidae). Annals of the Entomological Society of America, 100(6): 773-786. PDF

2007b.

Hall, J. P. W. Two new genera in the Nymphidiina (Lepidoptera: Riodinidae: Nymphidiini). Zootaxa, 1415: 35-42. PDF

2007c.

Hall, J. P. W., and D. J. Harvey. Three new species in the riodinid tribe Symmachiini from Panama and Colombia (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 7-10. PDF

2007d.

Hall, J. P. W., and G. Lamas. Four new symmachiine taxa from the eastern Andes of Peru and Ecuador (Lepidoptera: Riodinidae). Zootaxa, 1533: 41-52. PDF

2007e.

Hall, J. P. W., and K. R. Willmott. Four new species of Symmachiini from Ecuador (Lepidoptera: Riodinidae). Tropical Lepidoptera, 16(1-2): 1-5. PDF

2008.

Hall, J. P. W. Theope revisited: a synopsis of new discoveries, with the description of three new species from Ecuador and Peru (Lepidoptera: Riodinidae: Nymphidiini). Proceedings of the Entomological Society of Washington, 110(1): 144-158.

2009a.

Willmott, K. W., A. V. L. Freitas, J. P. W. Hall, K. L. Silva-Brandão, and M. Paluch. A new species of Actinote Hübner from the eastern Andes of Ecuador (Lepidoptera: Nymphalidae: Heliconiinae). Proceedings of the Entomological Society of Washington, 111(1): 47-56. PDF [NOTE: published figures are monochrome]

2009b.

Hall, J. P. W., and K. R. Willmott. Two new species of Euselasia (Riodinidae: Euselasiinae) from western Ecuador. Tropical Lepidoptera Research, 18(1): in press.

2009c.

Hall, J. P. W., and K. R. Willmott. Discovery of a new Lucillella species (Riodinidae: Symmachiini) in the eastern Andes of Ecuador using the single rope canopy access technique. Journal of the Lepidopterists' Society, submitted.

2009d.

Janzen, D. H. et al. (46 authors). Integration of DNA barcoding into an ongoing inventory of complex tropical biodiversity. Molecular Ecology Resources, in press.


2010.

Cock, M. J. W., and J. P. W. Hall. An annotated checklist of the Riodinidae (Lepidoptera) of Trinidad and Tobago. [in prep.]

Popular publications:
2001h.

Hall, J. P. W., and K. R. Willmott. Ecuador. Out on a limb. National GeographicMagazine (Update from the field), 199(2): 86 PDF

2001i.

Willmott, K. R., and J. P. W. Hall. Winged gems of Ecuador. Biodiversity paradise. National GeographicMagazine (Geographica) 199(5): xxviii. PDF

2003c.

Willmott, K. R., and J. P. W. Hall. Common Butterflies of the Ecuadorian Amazon. Published by the authors, London, UK. 2 pp.

2003d.

Willmott, K. R., and J. P. W. Hall. Common Butterflies of Yuturi and Yarina lodges. Published by the authors, London, UK. 2 pp.

2006c.

Eliot, J. L., and J. P. W. Hall. Fossils. Butterfly jewels. National Geographic Magazine, 210(1): xxx.